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Emilie Bess and Kevin P. Johnson
Containing group: Epipsocetae


Cladiopsocidae includes two genera.  The genus Cladiopsocus (16 species) is known from Angola and from Central and South America.  Spurostigma (11 species) is known from Central and South America.

Family Monophyly

Recent morphological studies of the family Cladiopsocidae suggest that it is not monophyletic (Yoshizawa 2002, Casasola-Gonzalez 2006).  Morphological study by Casasola-Gonzalez (2006) indicate that the genera Spurostigma and Cladiopsocus belong in distinct families: Spurostigmatidae and Cladiopsocidae.

Discussion of Phylogenetic Relationships

The genera Spurostigma and Cladiopsocus have not yet been included in molecular studies.


Casasola González, J.A. 2006. Phylogenetic relationships of the genera of Epipsocetae Psocoptera: Psocomorpha). Zootaxa 1194:1–32.

Eertmoed, G.E. 1973. The phenetic relationships of the Epipsocetae (Psocoptera): the higher taxa and the species of two new families. Transactions of the American Entomological Society, 99: 373–414.

Eertmoed, G.E. 1986. The redefinition of Cladiopsocus (Psocoptera: Cladiopsocidae). Studies on Neotropical Fauna and Environment, 21: 207–229.

Lienhard, C. and C. N Smithers. 2002. Psocoptera (Insecta) World Catalogue and Bibliography. Muséum d'Histoire Naturelle, Geneva, Switzerland.

Smithers, C.N. & Thornton, I.W.B. 1977. A new genus and some new species of Epipsocidae (Psocoptera) from the Melanesian arc. Proceeding of the Linnean Society of New South Wales, 102: 60–75.

Yoshizawa, K. 2002. Phylogeny and higher classification of suborder Psocomorpha (Insecta: Psocodea:'Psocoptera'). Zoological Journal of the Linnean Society 136: 371-400.

About This Page

Emilie Bess
Illinois Natural History Survey, Champaign, Illinois, USA

Kevin P. Johnson
Illinois Natural History Survey, Champaign, Illinois, USA

Correspondence regarding this page should be directed to Emilie Bess at and Kevin P. Johnson at

All Rights Reserved.

Citing this page:

Bess, Emilie and Kevin P. Johnson. 2009. Cladiopsocidae. Version 25 March 2009 (under construction). http://tolweb.org/Cladiopsocidae/14464/2009.03.25 in The Tree of Life Web Project, http://tolweb.org/

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